Front view of flower with red nectar in wells at base of corolla. Anthers have dechisced. Stigma next to dehisced anthers looks like it has pollen on it. Calyx hidden by corolla. Corolla has thickenings extending from androecium toward corolla lobules; these create pockets that hold nectar between them (Photo by Mione, Mione et al. 790)	Flower in side view; both corolla lobes and lobules (alternating with each other) are evident at bottom of photo, and shape of calyx lobe is shown at top center (Photo by Mione, Mione et al. 790)
Flower Bud. Calyx wide open, corolla closed (Photo by Mione, Mione et al. 790).	Back of calyx. Where two sepals meet there is a keel that protrudes opposite the direction of the corolla, and that pedicel is angular in cross section (not terete). (Smallest units on ruler are mm, Photo by Mione, Mione et al. 783).
Gynoecium and calyx (corolla with attached androecium had already dropped). Disk around base of ovary is orange. Stigma at top of style is darker than style. Four calyx lobes are evident and the 5th is out of focus and closest to viewer. (Photo by Mione, Mione et al. 783)	Jaltomata pallascana plants growing in/on rock wall at or very near type locality (Photo by Mione, Mione et al. 783, Segundo Leiva G. next to plants).
Stems of intermediate age (Photo by Segundo Leiva G., Mione et al. 783)	Stamen as viewed from gynoecium (ventral view). Note filaments insert on the ventral face of the anther as in all of the members of this genus. Hairs are purple, unbranched and grow only at the base of the filament (Photo by Mione, Mione et al. 783)
Oldest stems (Photo by Segundo Leiva G., Leiva 4429 = Mione et al. 783)	Corolla "C." White arrows point to the mucro on the end of the anthers, not present in all populations of this species studied in northern Ancash. Anthers "A" are undehsiced and yellow with purple dots; the lowest anther shows, toward viewer, a line where longitudinal dehiscence takes place. Filament "F." Stigma, darker green that the style, is immediately to the left of "S" (Photo by Mione, Mione et al. 783).
View across valley from plants of Jaltomata pallascana. (Photo by Mione, at or very near type locality, Mione et al. 783).	Ripe Fruits. One fruit collected in Peru was 21.5 to 22 mm (depending on how you rotate the berry) by 15 mm (pedicel attachment to opposite pole) and contained 301 seeds (Photo by Mione, Mione et al. 790).

Discussion: Georg Bitter described a new subspecies: Saracha weberbauer Dammer subspecies pallascana. Mione studied Bitter’s (1924) paper and determined that the described species is a Jaltomata, not a Saracha. To confirm this, in May of 2008 Mione, Leiva G. and Yacher went to the type locality, in the difficult to reach northern edge of the department of Ancash. We are apparently the first botanists to collect this species since the type specimen was collected 88 years ago, and the first to photograph the large green flowers having bright red nectar (photos above and described in the protologue). This taxon has been poorly known in part because the holotype was destroyed in Berlin during World War II. Based on our field studies we conclude that it was correct for Macbride (1930) to have treated this taxon as a species. However, it was incorrect for Macbride (1962) to have placed this taxon in synonymy with S. weberbaueri Dammer, the latter now Jaltomata weberbaueri (Dammer) Mione. Mione and Coe (1992), following Macbride (1962), incorrectly placed S. weberbaueri Dammer subsp. pallascana Bitter in synonymy with Jaltomata weberbaueri (Dammer) Mione. Alvarez (1996) correctly listed this taxon among those excluded from the genus Saracha.

Variation Seen Within The department Ancash, Peru: The corolla can look 10-lobed (lobes and lobules alternating) as in the top-right photo above, or the corolla can look 5-lobed (pistillate phase flowers of population 784), and collection 791 was described as 5-lobed in the field but the same flowers studied here with a dissecting microscope were considered weakly 10-lobed. Anthers were mucronulate in populations 783 and 784, but a mucro was lacking on the anthers of populations 790 and 791. Purple dots were seen on the yellow anthers of populations 783 and 791 (showing in photo above), but these were lacking in populaion 784 (data not available for population 790). Leaf size varies markedly among these collections as well: 783 has small leaves while 784 (just a few km away) has large leaves.

Taxonomy: Jaltomata pallascana (Bitter) Mione, Arnaldoa 15(2): 285-288, 2008. Saracha weberbaueri Dammer subspecies pallascana Bitter, Repertorium Specierum Novarum Regni Vegetabilis 19: 267-269. 1924. TYPE: Peru, Department Ancash, province Pallasca, in the valley of Rio Pelagatos, in an area with bushes alternating with patches of grass, not common, 3,600 - 3,700 m, 29 Mar 1920, Weberbauer 7225 (B, destroyed, photos at NY & WIS, F neg. 2559; Isotypes F, US). Saracha pallascana (Bitter) Macbride. Field Museum of Natural History, Botanical Series 8: 110. 1930.

Specimens Examined:

Plants of Jaltomata pallascana are immediately above Segundo Leiva's head. (At or very near type locality, Photo by Mione, Mione et al. 783).	Lago Pellagatos, a few km up (higher than) collection locality 783.

Literature Cited